
Lynn Firth Virtual Wildflower Walk
Trolley Line Trail, Erie National Wildlife Refuge
Welcome to the virtual tour!
Join us on for a virtual hike along the Erie National Wildlife Refuge's Trolley Line Trail in search of wildflowers and other signs of spring! This event is in loving memory of Foundation for Sustainable Forests (FSF) co-founder Lynn Firth, who outshone us all with her knowledge and appreciation of spring wildflowers.
Co-hosted by the Foundation for Sustainable Forests, PA District Forester Cecile Stelter, and Erie National Wildlife Refuge Biologist Melissa Althouse, we will share videos, photos, and information about the flowers along the trail via social media and the Foundation's website . Though we usually invite you to join us out in the woods for this walk, the threat of COVID-19 has forced us to think creatively. Enjoy your armchair visit to the forests of northwestern Pennsylvania!
About your guides
Melissa Althouse, Wildlife Biologist at Erie National Wildlife Refuge
Melissa graduated from Green Mountain College with a B.A. in Biology in 2009 and the State University of New York College of Environmental Science and Forestry with a MS in Wildlife Biology and Conservation Management in 2016. She has been with the Refuge for three years, and in that short time has initiated a number of partner projects such as the French Creek Cooperative Weed Management Area and a local chapter of the PA Native Plant Society. Her love for the outdoors is driven by a curiosity to know what everything is and how it connects to the larger ecosystem. She is particularly fond of birds and wetland plants.
Cecile Stelter, District Forester for the PA Bureau of Forestry
Cecile Stelter graduated from Penn State University in 1989 with a B.S. in Forest Science and in 1990 with a MFR in Forest Management. Cecile has worked for the DCNR, Bureau of Forestry for 20 years and has been employed as the District Forester since 2008, working in the Cornplanter Forest District based in Warren, PA. Cecile is a Life Member of the Pennsylvania Forestry Association and the Pennsylvania Forest Fire Museum Association. She is a Certified Forester with the Society of American Foresters and a member of the Plateau Chapter of SAF. She is also a Certified Arborist with the International Society of Arboriculture and a lifetime member of the Girl Scout organization.
Anne Maloney, Executive Director of the Foundation for Sustainable Forests
Annie Maloney is the Executive Director of the Foundation for Sustainable Forests. Annie holds a Ph.D. in Biology from Boston University, where her graduate research focused on understanding how interactions between climate change and pollutants affect ecosystem nutrient cycling and long-term resiliency of forests in the northeastern United States. She is an Adjunct Professor at Allegheny College in the Department of Environmental Science & Sustainability and a former wilderness guide for the Voyageur Outward Bound School in Minnesota.
About the trail
Interactive map of the Trolley Line Trail
Trolley Line Trail is a 1.9-mile one-way trail in the Seneca Division of Erie National Wildlife Refuge, about 5 miles east of the borough of Cambridge Springs. Wildflowers and historic bridge foundations are highlights of the primitive trail, which follows an old railroad track. This is the most challenging trail on the Refuge with a short, steep incline on the western side, but majestic forest views reward the effort.
Starting out at the Trolley Line trailhead
Guy Dunkle of the Foundation for Sustainable Forests cameos in our tour, and shares a bit of Trolley Line history
A wooden bridge guides hikers over meandering streams in the Seneca Division.
A typical scene showing the forest along Trolley Line waking up for Spring.
A steep section of the trail near the Swamp Road trailhead.
With all of those creeks and wetlands near the trail, plan to encounter some mud on your hike.
Wildflowers of the trail
You'll come across dozens of interesting plant species along Trolley Line Trail. Some of the more common wildflowers are in the following slideshow. Each flower has a small map showing a specific location where Annie has found it along the trail, in case you'd like to adventure out on your own later.
Many of the slides have multiple images to show how these flowers mature through the season, or what sort of habitat you'll find them in. Be sure to hover your mouse over the photos to display navigation arrows and scroll through.
Annie's favorite spot for wildflowers on the trail (slides 12-14 in the slideshow below).
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1
Skunk cabbage
Symplocarpus foetidus
Melissa: Though it's not a true flower, skunk cabbage is one of the very first things you'll see growing in Spring wetlands. Often times, you'll smell this plant before you spot it - it's very appropriately named! In addition to being stinky, it also generates it's own heat - if it's caught in a late Spring snow, you'll see a little clear circle around the blooms. These rank blooms fade by May and large, broad leaves expand quickly after.
One of the buds has been chewed off in the first picture. Wonder what might have a taste for something so rank? Skunk cabbage has mild laxative properties. Black bear will often graze on skunk cabbage when they wake up after their winter torpor so they can 'unplug.'
2
Spring beauty
Claytonia virginica
Cecile: One of the earliest and smallest blooming wildflowers of Spring, growing just over 6 in. tall. Stem contains one pair of long, narrow leaves. Flowers range in color from white or light pink to darker pink/purple and flower petals have distinctive dark pink lines running lengthwise along the flower petals. Found in early Spring in drier wooded sites. It is part of the Portulaca Family.
3
Violet
Viola spp.
Cecile: There are many different varieties of this common wildflower. Blue or purple are the most common species but there are also white and yellow varieties. The leaves are heart-shaped, and the solitary flowers are produced on leafless, individual stems. Violets usually bloom in May and do best in moist, shady sites but can adapt to other settings. Violets are one of the larval food sources for fritillary butterflies.
Scroll through the Violet photos to see an example of the three colors you can find in Northwestern Pennsylvania.
4
Spotted geranium
Geranium maculatum
Melissa: This showy relative of your garden geranium is often found in clumps in the woods or on the edge of fields. The 'spotted' in the name refers to the white specks on the leaves. Plants grow 1-3 ft. tall, with 1.5 in. flowers growing in small clusters above a pair of deeply-lobed leaves. Spotted geraniums are pretty easygoing as far as woodland wildflowers go - they'll grow in anything from full sun to dappled shade,and in moist hummusy soil to well-drained, poor soil. The nectar and pollen of this flower is an important springtime food source for our native pollinators.
5
Marsh marigold
Caltha palustris
Melissa: You can find carpets of these bright mounding flowers in Spring wetlands. Marsh marigolds can grow between 8-24 in. tall. They have fairly round, lightly scalloped basal leaves. Plant stems are hollow (a trait that helps differentiate it from a similar looking invasive species). The flowers bloom in clusters on a plant, average 1 in. across, and, though they appear to be just yellow to us, bees see them as being very colorful. To insects, the upper part of the petals appears as a mixture of yellow and an ultraviolet color, "bee's purple", while the lower part is all yellow. Unlike most of the other wildflowers in this slideshow, you'll usually find Marsh marigold in sunny locations.
6
Purple cress
Cardamine spp.
Melissa: Probably the prettiest plants in the mustard family (Brassicaeae), Cardamine douglassii (Purple Cress), shown here, is similar to Cardamine bulbosa (Spring Cress), except that the latter has flowers with white petals and green sepals and lacks a hairy stem. This wildflower blooms in the early Spring beneath deciduous trees with evenly moist soil.
Note that this is the place where Annie recorded the mud PSA - be prepared for some wet toes!
7
Trout lily
Erythronium americanum
Melissa: This flower is named for the mottled fish that the leaf pattern resembles. Though usually yellow, you'll occasionally come across a rare patch of white Trout lilies. It takes up to 5 years for a plant to start flowering (many species in the lily family take a long time to mature). This is one of the most common wildflowers you'll see in our woods. Trout lilies cameo in many of the photos for other wildflowers in this slideshow - see how many you can find.
8
False hellebore
Veratrum viride
Melissa: Also known as Indian poke and Corn-lily, this is a unique-looking plant found on floodplains and shallow wetlands. The large, deeply ribbed leaves are the prominent feature of this plant. False hellebore might be the largest plant in this tour - it can grow up to 6 ft. tall, though it's usually closer to 3-4 ft. It has a subtle cluster of star-shaped, hairy flowers that grows from a central stalk. The flowers fade quickly, while the leaves persist through early summer.
9
Slender toothwort
Cardamine angustata
Melissa: You can distinguish this species from other toothworts by looking at the leaves - Cutleaf has deeply incised leaves; Forkleaf has pencil-thin, delicate leaves; and Slender has variable leaves with a pair of broad basal leaves and a narrow pair that subtends a cluster of nodding white blooms. Plants will grow roughly 8-12 in. tall. Our flowers are just about to bloom - keep an eye out for these if you walk the trail closer to May.
10
Blue cohosh
Caulophyllum thalictroides
Cecile: Spreading and bushy when mature, blue cohosh is a large, multi-stemmed perennial, 1-3 ft. tall. Thrice-compound leaves with lobed leaflets are purplish in spring and become green as they develop. Inconspicuous, purplish-brown to yellow-green clusters of flowers are followed by conspicuous, bright-blue berries.
Be sure to scroll through the photos of Blue cohosh to see what it looks like as the seasons progress. The first photo was taken on April 22nd, 2020, to give you an idea of what to look for this time of year.
11
Red trillium
Trillium erectum
Melissa: This is my favorite wildflower! The plants grow from an underground rhizome, from which it takes years for a plant to mature and start blooming. Fun fact: Trillium plants produce no true leaves or stems above ground. The “stem” is actually just an extension of the horizontal rhizome and produces special leaves called cataphylls. These highly modified leaves surround the flowering scape (the above ground plant) as it pushes up through soil in early spring. The leaf-like structures are technically bracts subtending the flower. As the name suggests, all parts of this plant tend to be arranged in threes (the"leaves", flower petals, and reproductive parts), which is typical of plants in the Lily family (Liliaceae). The Red trillium is sometimes called a 'Wakerobin' as it blooms for just 2-3 short weeks around when the American robin starts to sing. You can find this plant in the understory of rich, moist woods.
Be sure to scroll to the 3rd and 4th photos to see another similar species, the white trillium (Trillium grandiflorum).
12
Wild ginger
Asarum canadense
Annie: Wild ginger occurs as a low, colony-forming perennial typically 4-8” high, primarily in the understory of deciduous forests in moist, rich soils. An individual plan consists of a pair of velvety, heart-shaped leaves. The solitary flower is often overlooked, hidden as it is at the base of the leaves, and is red or purplish in color. The rootstock does have a strong, gingery flavor, but is not related to commercial ginger (Zingiber officinale). Although used by early Americans as a seasoning, the plant is not typically used for culinary purposes, and in fact is considered toxic by the USDA.
This flower is part of the wildflower hotspot that Annie referred to in the video before this slideshow. It occurs along a stream, supporting the moist, rich soils that it needs to thrive.
13
Bloodroot
Sanguinaria canadensis
Melissa: This poppy relative is a true Spring ephemeral, with flowers that only last about 1-2 days. A solitary 6-10 in. bloom emerges from the rhizome wrapped up in a deeply scalloped leaf. Bloodroot flowers open during the day and close up at night. The leaves will keep developing over the summer after the flower has faded, and can reach up to 10 in. across. Colonies will grow together in the rich soils of streamside forests. The flower is named for the red sap present in all parts of the plant.
Also note that this spot on the map is the wildflower hotspot that Annie referred to in the video before this slideshow.
14
Mayapple
Podophyllum peltatum
Melissa: This is a good flower to practice on if you're just starting to learn Spring wildflowers. It's fairly tall (growing up to 20 in.) with large, conspicuous leaves. These plants are appropriately named, as they bloom like clockwork around early May in our region. A fleshy fruit that resembles an apple will follow later in the summer. Much like the Canada mayflower that you'll see elsewhere in the slideshow, Mayapple flowers only bloom on fertile shoots with two leaves. The infertile stems with just a single leaf will vastly outnumber the blooming ones within a colony. These plants make me think of little fairy umbrellas.
15
Canada mayflower
Maianthemum canadense
Melissa: One of the most common wildflowers you'll see out in our woods in the Spring. These diminutive plants grow about 4 in. high in dense clumps - but you'll only see the blooms on the plants with two or more basal leaves. The infertile single-leaf shoots usually outnumber the fertile blooms. These plants thrive in dappled sunlight with high humidity and cool temperatures. The berries, which form later in the year, are a favorite food source for Ruffed grouse (Bonasa umbellus). You'll often see mayflowers alongside Spring beauty and Trout lily.
16
Cutleaf toothwort
Cardamine concatenata
Melissa: You'll usually find this plant along the edges of small streams. The central flowering stalk reaches about 10 in. tall. It can be a bit weak and flop over, but will grow erect and the flowers will open more on warm Spring days. The leaves are deeply divided and coarsely toothed. You might think this is what gave it the common name of Cutleaf, but it's actually named for tooth-like projections present on the underground portion of the stem. Cutleaf toothwort needs undisturbed soils - its presence usually means that the immediate area has never been plowed under, or subject to vehicular or other forms of traffic.
Special wildflower spotlight: Cecile talks about Bloodroot.
Other wildflowers and wildlife in the area
Northwestern Pennsylvania has a rich, diverse array of natural resources, which you can find beyond the boundaries of Erie National Wildlife Refuge. Trolley Line Trail is an exemplary northern hardwood forest, but only certain plants grow in that habitat. Check out some of the other plants and wildlife that you'll come across in other natural areas.
Liverleaf
Hepatica nobilis var. obtusa
Cecile: One of the earliest, showy wildflowers of the forest. Multiple, thick leaves on hairy stems, that are heart-shaped at the base and have three deep rounded lobes, that often form clumps out of which the flowers emerge. Flowers may be bright blue, pink or white, up to 1 in. wide. They open their fullest on sunny days and have a fresh, delicate scent.
Red-spotted newt
Notophthalmus viridescens
Melissa: These newts are very common, and blend in surprisingly well with the leaf litter despite their bright colors. Watch your step as you hike along the trail! You'll often hear us calling these guys 'red efts' rather than by their true name - an 'eft' is the terrestrial juvenile stage of this otherwise aquatic amphibian.
Annie: Even the red efts are practicing safe social distancing! This guy was hiding in a rotting log when I walked by.
Ramps
Allium tricoccum
Melissa: Also known as wild leeks, these plants smell faintly of garlic. Large patches pop up beneath dry forests early in the Spring. Though some folks like to collect these plants for consumption, please remember that it is illegal to remove any plant, animals, or mineral from federally protected lands. Resist the urge to nibble while you're on the trail - it's a finable offense! If you can find a good patch on State Game Lands, though, you're welcome to some of the bounty.
Dwarf ginseng
Panax trifolius
Cecile: Diminutive plant 2-8 in. tall with 3-5 leaflets in a small whorl. Produces a rounded cluster or ‘ball’ of tiny white flowers in April/May. Found in moist woods. This tiny plant is related to the commercial Ginseng plant that is harvested for it’s root, but the roots of this small plant do not ever grow large enough to make it worth digging.
Jack-in-the-pulpit
Arisaema triphyllum
Cecile: A 1-2 ft. tall striking plant in which three leaves in a single whorl, overtop a ‘spathe’ or flower that resembles a “tube with a pointed hood.” The coloration of the flowers may be quite variable and can range from all green to purple or green with purple stripes. This plant grows in moist, rich, shady hardwood forests often alongside Trillium and flowers in late April and May.
Melissa: You can tell non-flowering Trillium and Jack-in-the-pulpit plants apart by looking at the leaf arrangements - Trillium's leaves are arranged in a radial whorl, whereas Jack-in-the-pulpit's leaves have a sort of flat edge on the bottom, where the lower two leaflets are opposite each other.
Fiddleheads
Melissa: Even plants that don't flower have something to show off as they emerge in the Spring. Here we see the fiddleheads of a fern unfurling.
Pink lady slipper
Cypripedium acaule
Cecile: A member of the Lily Family, the Pink Lady’s Slipper is one the largest native orchids, growing up to 18 in. tall. It produces two opposite basal leaves with conspicuous parallel veins and a large pink or whitish-pink flower at the end of a long, leafless stem. Flowers are produced in mid to late May or early June and plants can grow individually or in clumps. It is very site-specific and propagates poorly and is very difficult to transplant. Also called the Pink Moccasin plant.
Fossils
Melissa: A small fossil Annie found along the trail. Are fossils a passion of yours? Check out this guide to the Common Fossils of Pennsylvania . The resource might be a little old (1999), but with fossils being even older I think you'll still find it interesting. This one is a brachiopod. The official Pennsylvania state fossil is a species of trilobite. Remember, though finds like these are fascinating, leave them behind for others to also appreciate.
Spring peepers
Melissa: Listen to one of the ultimate sounds of Spring in northwestern Pennsylvania - a Spring peepers putting on his show! Despite their impressive chorus, Spring peepers are one of our smallest tree frogs. They only grow to a size of 1 in. - barely larger than a paperclip.
Vernal pools
Annie talks about the vernal pools visible alongside the trail, and common throughout the region.
Spotted salamander
Ambystoma maculatum
Melissa: Spotted salamanders are one of the Spring-breeding amphibians that depend on vernal pools for safe breeding habitat. Along with Wood frogs (Rana sylvatica), Spotted salamanders migrate from upland forests to low-lying ephemeral pools during warm, wet evenings after the snow melts. What makes vernal pools so attractive? As they are temporary bodies of water, fish can't live there - which means fewer predators to endanger the larvae/tadpoles that hatch out.
Four-toed salamander eggs
Hemidactylium scutatum
Melissa: Other amphibians are also breeding in the damp Spring atmosphere. This mass of Four-toed salamander eggs was found just upland of a vernal pool. You can start to tell the species of amphibian egg by looking at the embryo: these are two-toned and round, don't have much of an outer gel matrix, and were laid on a mossy substrate. All characteristic of the Four-toed salamander.
Wildlife viewing opportunities
Melissa: Snags, or dead trees, are common in mature forests and provide tons of wildlife value - from nesting cavities, to foraging opportunities in the decaying wood. This one was visible from the Trail near the Swamp Road trailhead.
Birdsong
Melissa: Spring is a time of life. Wildlife is often more visible as creatures display to attract a mate. Here, we see three eggs from an American robin (Turdus migratorius) safely tucked away in a grass-lined nest.
Morning display
Annie took this short video on her hike along the trail. You can hear the birds singing a bright morning chorus, and see a male blackbird displaying from the top of a snag nearby.
Sing it!
Melissa: This Dark-eyed junco (Junco hyemalis) was singing his heart out by the Refuge's Visitor Center last Spring.
We hope you enjoyed your visit!
Take care & thanks for joining us!
We welcome visitors to check out the trail on their own and see these beauties in person. Refuge trails are open from 1/2 hour before sunrise to sunset. Please stay on the trail and leave the flowers in place for others to discover. Leashed dogs are welcome to join you on your hike. Remember to follow CDC guidelines to protect yourself and others when enjoying the great outdoors.
- If you are sick, stay home.
- Keep a social distance (6+ feet) from others.
- Announce your presence to others.
- Stay regional - don't travel far from your residence.
- Avoid times and places of high use.
- Practice good hand hygiene.
- Be courteous and kind.