Fen Resilience in Southern Michigan
Mitchell's Satyr Response to Research & Habitat Management
Mitchell's satyr butterfly observed in southern Michigan during Michigan Natural Features Inventory annual surveys in 2021.
Butterflies attract our attention in a way few other animals can. This is the story about the efforts of scientists, land managers, and the public to protect a valuable population of one endangered butterfly species. This project represents a close partnership between the Southwest Michigan Land Conservancy (SWMLC) and Michigan Natural Features Inventory (MNFI).
SWMLC has been a local leader for over 30 years in the conservation and restoration of natural areas in southwest Michigan, including managing vital habitat for the Mitchell's satyr butterfly. MNFI collaborates with SWMLC and other conservation partners by providing scientific expertise for the management of rare species in Michigan. MNFI conducts research, including annual population monitoring, which informs land management.
Funding to support land management activities and population monitoring at Branch County Fen has come from the U.S. Fish and Wildlife Service (USFWS), Great Lakes Restoration Initiative, as well as essential community support from SWMLC members. The Toledo Zoo has been a partner at Branch County Fen as well, contributing to Mitchell's satyr captive rearing efforts.
This fen is not open to the public. For information on visiting SWMLC's public preserves, visit their website .
What makes Branch County Fen so special?
Branch County Fen contains a rich wetland complex that hosts dozens of rare species, preserves water quality in headwater streams, and represents a textbook example of a prairie fen natural community. Currently 39 acres of this fen complex have been protected by SWMLC through conservation work over the past three decades. Portions of the fen complex and adjacent uplands are owned by private individuals, who work with MNFI and SWMLC to manage their properties.
This wetland complex displays a range of natural conditions found in fens, including cold groundwater springs, species-rich sedge meadows with scattered native shrubs, and dappled shade under tamarack trees. The fen also contains an abundance of "marl flats" with soils rich in calcium and magnesium, where uniquely adapted plants like carnivorous pitcher plants (Sarracenia purpurea) and sundews (Drosera spp.) can thrive.
Fens are often degraded through one or more of the following: altered hydrology, water pollution, overgrazing, invasive plants, and fire suppression. Without management, these communities will often become either closed canopy shrub-carr or monocultures of narrow leaved cattail (Typha angustifolia) or other invasive species.
The Mitchell's satyr (Neonympha mitchellii mitchellii) is one of the most geographically restricted and critically endangered butterflies in North America. In Michigan, this small butterfly lives exclusively in fens where its host sedges (Carex spp.) are abundant.
Male and female butterflies are similar in appearance, however males are slightly smaller and darker than females.
Males emerge from chrysalides in late June and establish territories. Females emerge slightly later in the flight season, which typically ends in the second week of July.
A relatively sedentary butterfly, one study recorded daily movements of approximately 30 meters, with a maximum distance of 478 meters for females and 710 meters for males (Barton 2008).
Mitchell's satyr habitat goes beyond 'good fen'
Not all fens contain suitable habitat for the Mitchell's satyr butterfly. In addition to abundant narrow-leafed sedges, this species requires a shaded habitat component. Satyrs are often found within three meters of woody vegetation, such as native shrubs or tamarack (Larix laricina). Groundwater flow influences the vegetation patterning of these sites, and we are working to better understand the relationship between hydrology and Mitchell's satyr. Fens are fed primarily by groundwater, with hydrological input coming from a variety of sources, including springs, seeps, and lakes.
Partners from Michigan Natural Features Inventory, Fish & Wildlife Service, and The Nature Conservancy survey for Mitchell's satyr at a site in Jackson County, Michigan.
The bad news
Branch County Fen was initially in prime condition and supported hundreds of Mitchell's satyr butterflies. The butterflies could be found in pockets of sedge cover between tamaracks and poison sumac near groundwater springs. In recent years, the Mitchell's satyr population at this site has decreased, likely due to a combination of invasive species, woody encroachment, low genetic diversity, and hydrology changes. Below, we take you through the story of the discovery, monitoring, protection, and management of this site over time.

Number of Mitchell's satyrs documented at the fen from 2003 to 2022
Timeline
1965-1999
In 1965, Mitchell's satyr was first documented at Branch County Fen from a collected specimen. At this time the butterfly was not listed as endangered.
During a 1987 survey, no Mitchell's satyrs were found. Surveyors had possibly missed the narrow flight season by visiting too late in July.
In 1991, the Mitchell's satyr was added to the U.S. List of Endangered and Threatened Wildlife and Plants. That same year MNFI observed 34 butterflies during three site visits.
2000-2004
In 2000, MNFI received a USFWS grant to survey for the Mitchell's satyr across Michigan. At this point, there were over 20 known satyr sites. MNFI subcontracted with SWMLC to assist with monitoring at several sites in southwest Michigan, including Branch County Fen.
The Michigan Department of Environment, Great Lakes, and Energy observed a wetland violation by a private individual in habitat occupied by the Mitchell's satyr at Branch County Fen. Together with MNFI and USFWS, talks began with the landowner who had dug ponds and cut horse trails through the fen. Rather than paying a fine for the wetland violation, the neighbor worked with partners to protect the land. This resulted in SWMLC purchasing a 15-acre section of occupied habitat with funds provided by the USFWS.
Open sedge meadow adjacent to native cover in the oak upland was once grazed by cattle.
These 'rooms' of sedges surrounded by native shrubs and tamarack trees create ideal satyr habitat.
2005-2007: A new preserve!
After the land deal was finalized, the first 15-acre section of the fen was protected, shown by the yellow polygon on the map. SWMLC was now a landowner of an endangered species population. The Conservancy was able to start land management by working with Blue Heron Ministries , an Indiana-based non-profit.
Over the years, SWMLC and Blue Heron Ministries worked to create a 'corridor' to encourage butterflies to move between the core population and the southeastern pocket of the fen. This decision was fortuitous, as in later years this area would be a critically important habitat patch for the butterfly.
2008-2009
In 2008, regular monitoring of the site showed that the Mitchell's satyr population was dwindling.
In 2009, SWMLC conducted a prescribed burn (red shaded area on the map) with the goal to combat encroaching woody vegetation, which was shading out host plants needed by the Mitchell's satyr.
The prescribed fire was concentrated in areas where the butterfly was not found at the time. This was the first time in decades - if not centuries - that fire had returned to this landscape.
Fire helps create the open structure satyrs prefer by thinning shrubs and stimulating the sedges.
Patchy fire leaves unburned refugia for butterflies and other wildlife.
2010-2014
Population monitoring of the Mitchell's satyr continued, and showed that the butterfly had returned to the area burned in 2009.
Open fen habitat with shade from tamarack in the center portion of the preserve in 2012.
Invading cattails and shrubs became a problem in areas of main fen in 2013.
2015-2017
Mitchell's satyrs are found throughout most of the site, although their numbers are lower than in previous years.
Habitat work aimed to created a corridor to the southeastern portion of the fen continues with funding from the Michigan Department of Natural Resources.
2018
In a disappointing year, less than 15 butterflies were found during multiple surveys.
Notably, the southeastern fen pocket, which was once not suitable satyr habitat, now contained the core population. This area was targeted for future invasive species treatment and shrub/tree removal. This critical land management created excellent butterfly habitat which would not have existed without SWMLC action.
2019: A land deal!
Surveys continued to disappoint, with population contracting to an even smaller area in the southeast portion of the preserve.
By the time SWMLC acquired an additional 24 acres (broken yellow boundary) of occupied habitat and oak upland in 2019, the butterflies were restricted to just 13 satyrs in a pocket of habitat smaller than a football field. Buoyed with financial support from the USFWS, and guided by MNFI's technical understanding of the species, SWMLC began decisive habitat restoration to bring the species back from disappearing.
2019-2020: Winter work
Land management began in the winter of 2019-2020 as population decline led the satyr to become isolated in the southern portion. Using "surgical precision" SWMLC returned to formerly occupied habitat patches and removed just enough brush cover, while leaving important native plant species like bog birch, poison sumac, red osier dogwood, and tamaracks. Mitchell's satyr habitat preferences are often described as a "Goldilocks" scenario, with not too much shrub cover, but not too little, and just the right species of sedges near groundwater springs. The restoration crews attempted to "create" new habitat where Mitchell's satyrs hadn't been seen in years.
Winter is a great time for land managers to cut shrubs in prairie fen habitat to minimize habitat disturbance.
2020: Summer surveys
SWMLC and MNFI counted a small victory in the annual butterfly counts of 2020! Three Mitchell's satyr were seen in a 1/4 acre of previously unoccupied habitat that had been restored the previous winter! With enthusiasm renewed, a more extensive habitat restoration occurred in winter where crews "rolled out the red carpet" and tried to welcome butterflies into newly created habitat through use of an open flight corridor.
In 2020, mere months after shrubs were removed to re-create satyr habitat, we documented butterflies in this habitat for the first time since 2017.
2021: If you burn it, will they come?
In April SWMLC and Blue Heron Ministries conducted several small prescribed burns on habitat that hadn't seen fire for hundreds of years. The strategically positioned fire crept through 24 acres of prairie fen and oak upland to control woody vegetation and stimulate sedge growth to encourage the butterflies to expand into newly burned areas. The burns appeared to have a great impact on habitat, and in weeks new shoots of sedges were popping up through the wetland soils - but did the Mitchell's satyrs approve of the work?
When this extreme habitat makeover became available to adult butterflies the following July, surveyors were ecstatic to not only count roughly 40 butterflies in the core population, but also to find wandering male and female butterflies in the flight corridor cut AND over 300 yards away in corners of the preserve where they hadn't been seen in the past four years.
Prescribed fire helps to thin a brushy tamarack stand to create better habitat structure for satyrs.
A member of the burn crew monitors a burn break before spreading fire further.
The fire reveals the "hummocks" of a satyrs favorite food - Tussock sedge (Carex stricta).
Shortly after the burn the vegetation grows vigorously with nutrients returned to the soil.
2022
Mitchell's satyrs continue to reoccupy the northern, central, and southwestern portions of the wetland. Numbers were not incredibly high, however, with continued diligence and planning from land managers, the prognosis for the viablity of this population is not as dire as it once was.
This year we also released biocontrol beetles to knock back invasive purple loosestrife, which has invaded areas where invasive shrubs have been treated. In July we noted extensive feeding on the invasive wetland plant.
What comes next?
While the makings for a comeback story are present in the fen, we'd be mistaken to just kick up our feet and count butterflies. Hybrid cattails are present on-site and can be highly stimulated by prescribed burns. There is no rest for conservationists. Continued management of these species will be essential. SWMLC has secured funds to continue to manage invasive shrubs that have closed in on the preserve.
Check out our partnership with Querkus Creative to help conserve prairie fens by purchasing original artwork, including this Mitchell's satyr print:
Thank you to all of the partners who have dedicated their time to better understand and protect the Mitchell's satyr and its habitat in southern Michigan. Special thanks to Daria Hyde, Mitch Lettow, Dave Brown, Nate Fuller, and Nathan Hilbrands.