Virtual Mt. Lemmon Field Trip

Field Botany Laboratory (RNR 230L)

The Santa Catalina mountains provide amazing environmental diversity and opportunities for field botanical work

The Santa Catalinas are one of the "Sky Island" mountain ranges of the southwestern United States and northern Mexico. Because of their extreme changes in elevation, these ranges have remarkable environmental and biological diversity. The Catalinas rise more than 2100 m (6890 feet) above central Tucson and the University of Arizona campus. This virtual field trip will take you up the General Hitchcock Highway, which is also locally known as the Catalina Highway and the Mt. Lemmon Highway. It begins on the south (Tucson) side of the range and goes for about 48 km (30 miles) from the cacti and shrub-dominated arid environments at the base to the much moister temperate forests at the summit of Mt. Lemmon at an elevation of 2791 m. We will be talking about individual plants but also about how various species of plants occur together at a particular location, what is known as vegetation.

Lower elevation environments (left) are often dominated by cacti such as saguaro and drought-deciduous shrubs such as ocotillo, which is flowering in this image that was taken in March. Less than 45 minutes away by car is Bear Wallow (right), a moist temperate forest whose vegetation (all plants collectively) is more typical of the northern Rocky Mountains than of southern Arizona.

This field trip is best viewed on a laptop or desktop computer at full screen. Some of the maps and imagery will not display if viewed on a mobile device. Additionally, some web browsers do not support all of the graphics that are embedded in this field trip. If it doesn't work in your browser, try a different one (Chrome may be best, Firefox the next best, Safari the third best) and make sure you have the most recent version installed. There are times when the server where this file is located may be down. If this happens, just wait an hour and try again.


Climbing the mountain

We will begin with information on using contour maps and other geographical tools to visualize the environment. Then, as we proceed up the highway, we will stop at various points, and with images and explanation learn about plants there, how to identify them, and the environments that they inhabit, why this matters, and how they are changing. Certain terms are in bold text. These should become part of your botanical lexicons. You should record at least six of these terms (not including plant common names) and provide a definition for them in your Field Notebook. You may build your definition from information presented here, but you may also want to consult resources on the Internet. There is another exercise for you to complete within this trip, which you will also include in your Field Notebook.

There are lots of linked images and associated text. You should slowly explore both. Some buttons will lead to detailed descriptions of particular plant species. Take a close look at these, not with the goal of memorizing anything, but just to learn more about the amazing diversity of plants you can encounter in this mountain range and how they are described. A major goal here is to prepare yourself to better understand natural environments so that you can function effectively and enjoy them more when you're next in them.

Our focus will be mostly on larger, easier to see woody plant species. This is always a good starting point as you learn about plants in a particular area. Once you're on the ground, it's easier to then focus on the smaller, less conspicuous species. We will use common names for all organisms while on this field trip. A list of common names and their scientific names is available  here . (Some links will only be available to students registered in RNR 230L.)

Plants and the physical environment

The Catalina Highway quickly climbs along slopes and canyons with the grade averaging about 5-8% (5-8 m rise for every 100 m covered on the road). We will encounter many different landforms (a natural feature of the earth's surface) including arroyos, ridges, drainages, cliffs, marshes, canyons, streams, hills, and summits. In order to function effectively as field botanists, we need to visualize the landforms we work in and be able to accurately describe our location. This is especially important when making a plant collection and our work may require that we read and interpret contour (topographic) maps. We will prepare for our journey by learning about these maps and what they tell us about the natural environment using topography of Sabino Canyon on the south side of the Santa Catalinas as a platform.

Reading a contour map

Contour maps allow us to visualize slopes and changes in elevation in a two-dimensional presentation.

  • Every point along the same contour line has the same elevation.
  • The contour interval (distance between any two lines) is the same throughout the map.

    • Every fifth contour is a darker color and is called an index contour. For example, if the index contours are labeled at 250 m, 500 m, 750 m then the contour interval is 50 m.
  • Each contour map legend (or metadata for a digital map) will include the map units and scale.
  • Map units are units of measure, such as meters or feet (this map is in meters).
  • Map scale indicates the relationship between distance on the map and corresponding distance on the ground. For example, this map has a 1:15,000 scale, meaning that 1 m on the (printed) map is equal to 15,000 m on land, and 1 cm on map is equal to 15,000 cm (150 m) on the ground. (Note that the scale on this digital map cannot be used for navigation because the web browser resizes the map to fit different monitors.)
  • Contour lines do not cross each other.

What is the contour interval on this map?

Slopes and hills

  • Widely spaced contour lines indicate a gentle slope, closely spaced lines a steep slope. Lines that nearly touch indicate a cliff.
  • One side of a contour line is "uphill" and the other side is "downhill."
  • Concentric contour lines indicate a hill with the inner most circular shaped line representing the highest point or local peak.
  • A saddle is the gap between two local peaks where the land falls away on either side, much like the seat on horse's saddle between the horn and cantle (high points).

Water flow

Contour lines that cross a water course (wash, drainage, stream, creek, river, etc.) are bent in the shape of a "V" with water flowing out of the open end of the V (see map for examples). Said another way, the open end of the V faces downhill.

To find the direction of flow:

  • Look for V shapes on the contour map.
  • Determine the elevation at two consecutive contours.
  • Water will flow from the point end of the V at the higher elevation, through the open end of the V and down into the point end of the V at the next, lower elevation and so on.

Opposite of a water course is a ridge, which is represented by "V's" with the open end of the V facing uphill.

Can you identify other water courses and ridges on the map?

From contour lines to 3D

With practice, one can look at a contour map and see the three-dimensional (3D) representation of the landscape.

Here we have draped our two-dimensional (2D) contour map over a digital terrain model of the same area.

Click anywhere on the map and then use the tools on the bottom right of the map to navigate the 3D scene in any direction and perspective. Activate a tool and then click and drag with your mouse or track pad to maneuver the map. The circular arrow tool is a good one to begin your exploration with.

As you tilt and rotate the scene, follow the contour lines and then visualize what the environment might actually look like on the ground based on these representations. Where will water flow and accumulate? Where might it be easiest or most difficult to travel?

Desert bighorn sheep are found in this environment. They tend to prefer habitat along ridge lines over steep slopes. Can you identify any of these sites on the map?

Game camera image of a desert bighorn sheep on a steep slope in the Santa Catalinas. Photo: Brett Blum.

Do the contour lines make more sense now?

Do the features pop out? Can you find the water courses, ridges, peaks and saddles?

Let's link this with the landscape on the ground

Sabino Canyon

The large canyon we've been displaying is Sabino Canyon on the southern edge of the Santa Catalinas. (In the canyon, look for the darker green ribbon of vegetation along Sabino Creek.) Like most of the range, it is part of the Coronado National Forest and is one of the most popular outdoor attractions in Tucson.

A paved road runs up Sabino Canyon from a Visitor's Center at the base of the canyon. Trams carry visitors up the canyon stopping at various locations. Lots of folks also walk or ride bikes (at certain times) up and down the canyon. Many trails in the canyon are very popular, especially in cooler periods of the year.

Water flows in Sabino Creek are fed by springs and precipitation over a large watershed on the south side of the Santa Catalinas. Plants found near the creek, in what is called a riparian zone (along the banks of a watercourse), are often very different from those away from this zone.

The creek flows under and sometimes over the road. Flooding and the physical disturbances it causes are an important factor affecting riparian vegetation and the animals associated with it. Photos: Kandres Halbrook.

Upscale 2D Technology

Although 3D maps are amazing and very instructive, 2D contour maps are still valuable because they condense a tremendous amount of information into a compact, technology-free format. Take a printed contour map and a compass with you into the field and you can identify where you are, where you have been, and you will not have to worry about battery drain, cell signal strength, or satellite coverage.

However, this doesn't mean that we have to eschew technology.

GeoPDF and GeoTIFF

A GeoPDF/TIFF is a Portable Document File (PDF) or Tagged Image File Format (TIFF) that has digitally embedded geospatial location data (GPS coordinates). The GeoPDF/TIFF can be viewed on screen and printed as a regular PDF/TIFF, such as the portion of the USDA Forest Service Sabino Canyon Quadrangle GeoTIFF shown here. However, when used in conjunction with a mobile mapping app and a GPS-enabled smartphone or tablet, the geospatial embedded data are accessed and one's location can be plotted on the map, much like the blue orb one sees when using an app such as Google Maps.

Good for field work

For field botanists, the benefit of this technology is that one can use a GeoPDF/TIFF when in the field and have the GPS coordinates of their collection locations and transects plotted on the map. The apps use the mobile device's GPS so one doesn't need an Internet connection or cellular signal, which is a plus given that many remote or mountainous areas do not have adequate cell or satellite coverage.

Another benefit of this technology is that it can greatly simplify route finding. Have you ever been out in a wildland environment and not been exactly sure where you are? Using GeoPDF/TIFF maps will make this less likely to occur.

GeoTIFFs for the USA (with English units of measure) can be downloaded free of charge from the  US Geological Survey  and are an example of public funding to support science, commerce, transportation, urban and rural planning and other endeavors where accurate maps are vital.

Using a mobile mapping app is not required for this course, but it is something that some of you might be interested in. There are a number of geospatial mobile apps to choose from and if you would like to pursue this technology you can shop around for the one that suits your needs (and be certain to read the app's privacy statement). One place to get started is  Avenza Maps  because it is free for up to three maps. There are many video tutorials for using this and other apps on YouTube and other sources.

Climate and vegetation

Wherever you are on land in the world, climate is the primary factor determining vegetation that occurs there. Your experience has already convinced you of this. For example, you probably accept that the vegetation of the Amazon Basin is what it is because the environment there is consistently relatively wet and warm. Plants adapted to those sort of environments occur there. Similarly, deserts of the southwestern United States have vegetation that reflect the fact that it is relatively dry and warm in these environments. Climate is also the major factor driving the vegetation changes we’ll experience as we go up the Catalina Highway. And this climate changes very rapidly here since elevation does the same. Average annual precipitation is about 300 mm at Tucson (elevation 728 m) but it is 751 mm at Summerhaven (elevation 2500 m), which is near the summit. The image below shows the south face or southern aspect of the mountain range and predicted mean annual precipitation.

Predicted mean annual precipitation of the Santa Catalina Mountains and the surrounding area based on WorldClim data.

As anyone who has ever driven up the Catalina Highway from Tucson will tell you, temperature also changes with increasing elevation. Weather instruments tell us that the annual average temperature is about 12° C (21.6° F) higher in Tucson than Summerhaven. This is a reflection of a physical characteristic of the atmosphere called the lapse rate, which is the change in temperature with change in altitude. While meteorologists have many ways of defining the lapse rate, depending mostly on the amount of water in the atmosphere, a good average lapse rate is about 6.5° C per 1000 m. The difference in elevation between Tucson and Summerhaven is 1772 m, thus 1.772 m x 6.5° C per 1000 m = 11.5° C, which is very close to the actual average temperature difference between the two locations from the instrument record. Lapse rate can tell us a lot about how vegetation will change with changes in elevation, but again there are many other geographical features to consider when trying explain vegetation on the ground.

Predicted mean annual temperature of Santa Catalina Mountains and the surrounding area based on WorldClim data.

Traversing the plant community types in the Santa Catalinas

Sonoran Desert scrub

We begin our journey at the base of the range where the vegetation is similar to that of much of the city of Tucson. This is a relatively dry and warm environment. The plant community type here is often referred to as Sonoran Desert scrub. Cacti including saguaro are common as are shrubs like creosote bush and scattered perennial grasses. Annual grasses and broad-leafed species will grow following cool- and warm-season rains, although the species will differ between seasons.

How to navigate the map

Click anywhere on the map to begin interacting with it. As with our 3D contour map, above, this 3D scene can be tilted and rotated in any direction to help you visualize the landscape. At the upper right is a Layers and Legend panel. Click on and off the "check boxes" in the layers panel to isolate individual features and explore the attributes of the scene. Click on a camera icon to see an GPS-linked image taken at that location. Click on the thumbnail to see a larger version of the image.

Species information

Click on the buttons below, to see images and taxon descriptions for each of the highlighted species. The points on the map for each species represent locations where specimens have been collected and entered into a herbarium database. For species that are commonly observed, often there are few collections. We should interpret the locations as confirmations of suitable habitat for the species but not as an outline of their entire range. Note also that traditional plant collections often are geographically biased, that is, they more often occur near roads or trails (confirm this for yourself as you explore this map and the ones that follow). The saguaro locations are based on identification of individual plants within 1-km grids from satellite images. As such, they are not geographically biased.

Typical Sonoran Desert scrub plant community near the base of the Santa Catalinas. The darker green shrubby trees here are velvet mesquite. Photo: Steve Smith.

Drought-deciduous shrubs

Along the lower slopes in Sonoran Desert scrub we will also see many shrubs and small trees. Brittlebush is a drought-deciduous shrub that is prominent here, especially in drainages. In most years it produces broad leaves with fewer hairs in the winter and spring, and drops these to produce smaller much hairier leaves in the summer. Smaller leaves plus hairs that reflect light and slow air movement around the leaf will all reduce water loss from transpiration, which is important for survival through the warmer period of the year. The hairs make summer leaves appear a much lighter green--almost white sometimes. Desiccated winter leaves that have not yet dropped are visible along with the light-colored and expanding summer leaves (upper left). Broad and less hairy (and therefore greener) winter leaves are in the upper right.

The lower images were taken in April when brittlebush was in flower. Brittlebush will produce a fairly vibrant display of color in the springtime in most years, even when precipitation in last few months has been well below average.

Saguaro habitat and winter temperatures

Saguaros are quite common as we begin the climb. Suitable habitat for saguaro is determined primarily by summer-fall precipitation and temperatures, and perhaps more importantly, by winter temperatures. Saguaro seedlings less than one year old are especially vulnerable to freezing temperatures. It’s often observed that saguaros will not survive on sites where on average there is at least one day per year when the temperature does not go above freezing (0° C). As we proceed up the highway, turning northeast above Molino Canyon, fewer mature saguaros occur, even on the warmer south-facing slopes.

This map shows 1-km cells where at least one mature saguaro occurs. As the mean January temperature decreases below about 10˚C saguaros do not occur.

Roads may provide excellent habitat for weedy species

Any plant growing where we don't want it to be is considered a "weed." Roadways and their rights-of-way are significant disturbances and rainfall concentrators, and can provide excellent habitat for weedy plant species. In the first 8 km of our trip, we see many of these species, especially in drainages in this desert scrub environment. Prominent among these are the closely related perennial species, fountaingrass and buffelgrass. Both of these species are native to Africa and southern Asia and were introduced to southern Arizona in the mid 20th Century for use in landscapes (FG) and rangeland revegetation (BG). These drought-resistant species may grow in areas between woody or succulent plants in Sonoran Desert scrub and increase the chances for large-scale wildfires, which are naturally very rare in these plant communities. Land managers and other groups are actively working to reduce stands of these species using herbicides and manual removal.

Two large clumps of fountaingrass, one actively growing (left) and the other appearing to be dormant (August). This image was taken on the downhill (south) side of the Catalina Highway. The rocks occur here because of highway renovation that began in the 1980s. Many weedy species are well adapted to this sort of soil disturbance. Photos: Steve Smith.

The image at right was taken in August and shows multiple plants of fountaingrass growing below the Catalina Highway.

Desert grassland and oak woodlands are above Sonoran Desert scrub

Boundaries between plant community types are fluid

As we ascend and climate changes so will the vegetation. One community type will generally blend into another, often quite gradually. You can see that in the map at right as the light green color representing the Sonoran Desert scrub blends into the darker green representing a community type known as desert grassland that then blends into the light blue of the oak woodland community (as always, click on and off features within the layers to explore the map attributes).

We've talked a bit about how elevation (climate) is a major factor determining plant distribution, but aspect (the compass direction that a slope faces) is also very important, especially in relatively arid environments such as the Santa Catalinas where water is a major driver of plant establishment, growth, and survival. At a given elevation in the Northern Hemisphere, south-facing slopes will be on average warmer and drier than north-facing ones. Slope will also affect soil properties, which then affect water retention and plant community composition.

Desert grassland

As we climb up past Babad Do'ag above Molino Canyon, the plants we associate with the lower desert become less common and perennial grasses like sideoats grama begin to occur. Small, multi-stemmed ("shrubby") trees like velvet mesquite provide some overstory cover, but most of the vegetation is grass. This is the desert grassland community. Large areas of the Sonoran and Chihuahuan Deserts have this community type, especially south and east of Tucson into northern Sonora and southern New Mexico.

The light brown portions of the hillside--elevation 1350 m--are dormant grasses (August). Mature saguaros, although still present, are much less common at this elevation. Photo: Steve Smith.

Oak woodland

The vegetation of Molino Basin is also dominated by perennial grasses. However, sufficient precipitation occurs here to support more trees, specifically oaks. Oaks, which are in the genus Quercus and the family Fagaceae, are most easily identified by their production of acorns. This genus is taxonomically complicated as hybridization between what are considered different species is common. Thus, intermediate types (interspecific hybrids) are also common. In the Santa Catalina range there are seven species of oaks, many of which occur in Molino Basin. This community is referred to as an oak woodland, which means one in which trees are present, but at lower density (number of individuals per some unit of area) than we would see in a forest.

Sight identification is a key tool for field botanists

In this course we focus a lot on sight identification of plants to families. The same knowledge and set of procedures can be applied to the sight identification of individual species—something that you actually have some experience with already. If you are an Arizonan, you could probably sight identify a saguaro long before you took field botany! For natural historians, identifying individual species is often very important. We’ll learn later in this course that when confronted by an unknown plant species--one we can't sight identify--being able to sight identify the family will greatly simplify species identification in most cases.

Like all good natural history, sight identification begins with an understanding of the environment you find yourself in at any point in time. We start by answering a series of questions: What’s the plant community type and elevation? What is the slope and aspect? What soils are present, how deep are they, and is water nearby or will it accumulate? Are fires, droughts, floods, or windstorms common in this environment? What animals (including humans) and other plant species might be affecting the species you are interested in? Knowing these geographical and ecological details, and then considering them together, will greatly reduce the number of possible species we might expect a a given site.

With this narrowed list of species, we then look closely at the plants to further limit the possibilities and hopefully come to a conclusion. (We have produced an  extended checklist  for sight identification.) As we do this, realize that everyone will do sight identification slightly differently. Accumulating botanical knowledge and experience with plants will likely change how you do sight identification. We will go through some examples of sight identification at stops along our field trip, beginning with a prominent oak woodland species.

Mexican blue oak

This is the first of our sight identification examples: A tree with broad, stiff, and toothless leaves that are blue green above and paler below. Fruits are egg-shaped acorns with 1/3 in the cap-like cup. The bark is gray or whitish and fissured in small plates. Secondary and lower veins are not prominent (ridged) in the leaves.

The observation that acorns are present is critical as this is a defining trait for the genus Quercus. Knowing the appearances, preferred habitats, and elevation ranges of southern Arizona oaks would lead us to conclude that this is Mexican blue oak. It’s found at the upper edge of desert grasslands and in mid-elevation woodlands and canyons between about 1300 and 1650 m. This species is often browsed by deer and many small mammals consume acorns. Examining images and descriptions of this species from a reliable source such as  SEINet  is a good way to confirm your identification, although this requires an Internet connection.

Notice that doing sight identification requires very careful and through observation. Often at least two of your senses (sight and touch) are involved; sometimes smell and hearing are as well. Here, we're focusing on the most important of these senses, which is sight. Remember, that this field trip is giving you a chance to look at lots of images. With this, it's up to you to determine what you actually see the next time you are in the field. If you don't get the difference between these two, think about listening and how sometimes this is not the same as truly hearing.

Fire

Fire has played a significant role in the vegetation composition of much of the Santa Catalinas. Heading up Molino Basin, about 1 km north-northeast of the  Gordon Hirabayashi Campground  we can see evidence of the Aspen Fire, which burned here in 2003. Before that fire, this was an area with mostly scattered oaks (it is a woodland), a few alligator junipers, and a grass understory. This was what the site looked like in November 2003 about four months after the fire. Note that some basal re-sprouting is visible suggesting these trees are not dead. 

The Aspen Fire burned for about a month in 2003 over an area of about 34,000 ha and destroyed 340 structures in Summerhaven near the summit of Mt. Lemmon.

October 2016, 13 years after the Aspen fire. Photo: Google.

The image above shows what the site looked like in October 13 years later (2016). Many of the oaks that seemed dead in 2003 have recovered and new oak seedlings, as well as those of other perennial species have established. Vegetation recovery from wildfire may not involve conversion into a new community type, but it can occur. Many factors will determine what happens following a fire including the initial plant community type and intensity of the fire, the time since the last fire, and the environmental conditions following the fire, especially those affecting erosion and seedling establishment.

Heading toward Bear Canyon: revegetation can have unintended consequences

Construction on the Catalina Highway began in 1933 and was not completed until 1950. A series of major road renovation projects were conducted between 1988 and 2007. Roadside revegetation, which mostly involved seeding, occurred as part of these projects. The goal in this work was typically to stabilize the soil in areas disturbed by construction. This revegetation has resulted in some interesting botanical anomalies—usually plants growing near the roadway that would not have been expected at this site otherwise. For example, the grass, blue grama is commonly included in revegetation seed mixtures and now occurs frequently along the road above about 2200 m, but is rare elsewhere in the range.

Blue grama. Photo: Max Licher.

Contaminates in revegetation seed mixtures can also introduce weedy species that may have very detrimental ecological consequences. Cheatgrass is a fire-adapted non-native winter annual that was only observed occasionally in the Santa Catalinas before the late 1980s. It is now quite commonly observed, especially near the roadway.

Cheatgrass growing in rocks along roadside drainage of the Catalina Highway (April). Photo: Kandres Halbrook

Cheatgrass along the Catalina Highway near Bear Canyon (April). Photo: Kandres Halbrook.

Much of the seed used in revegetation is produced in arid environments in the Great Basin where cheatgrass is especially common, especially (Utah, Idaho, and eastern Oregon). Our observations along the Catalina Highway suggest that cheatgrass was a weed in fields whose plants produced the seed of preferred species in roadside revegetation, like blue grama. As field botanists, we must always be careful when drawing conclusions based on plants at the roadside: non-native species are common, including some that may be weedy, and water runoff and concentration can make the environment locally wetter.

The image at right is from Thimble Peak Vista on the Catalina Highway (March 2017). The foreground shows grasses, oaks, and numerous shrubs regenerating after the Aspen Fire. Thimble Peak is visible on the ridge in the center of the image. This area was also affected by the Bighorn Fire in 2020. Photo: Steve Smith.

More on aspect and slope

Below is a 3D representation of an aerial image of Bear Canyon with the Catalina Highway winding through the fore- and middle ground. The image has been rotated so that we can look up the canyon along the highway. The Middle Bear Parking Area (elev. 1791 m), which has rest rooms, is located at the "star."

Note the vegetation differences between the slopes to the right and left of the portion of the highway that runs along Bear Canyon, shown in red. Vegetation on the cooler, moister north-facing slope is both more abundant and of a different type than that on the south-facing slope to the left of highway.

An activity for you to do

As we often say in Arizona, "it's cooler in the shade." Have you ever noticed that it is blistering hot standing outside near a south-facing wall of a building, but much cooler on the north side of the building? Wherever you live, go outside and find, photograph, and describe a location that shows obvious differences in plant growth, health, or composition as a result of aspect. Places with topographic relief are good places to look, but <big hint> you probably don't have to go far from home, wherever your home might be. Include all of your observations in your Field Notebook, of course with a  header page  describing the location as always.

Image: Google Earth Pro 7.3.3.7786 (August 18, 2018), 3D Scene: ArcGIS® software by Esri

The image below is aspect raster (raster: digital image) of the aerial scene, above. The raster was created by analyzing 1 km x 1 km grids containing elevation data. Each color represents a compass direction that the slope faces. Although such fine-scale data manipulations are not necessary for day-to-day field botany work, this technology and others are regularly used by land and watershed managers, wildlife biologists, botanists, fire ecologists, and others to categorize landscapes in order to understand current conditions and predict future changes, such as how a landscape will respond to fire, and how changes in vegetation will affect watershed properties, or threatened and endangered plant and animal species. From the aspect raster, we see that the cool side of the canyon is predominately northwest facing and the opposite side is south-southeast facing.

Aspect raster created from a 1-arcsecond digital elevation model

Bear Canyon

A very cool spot

We next enter Bear Canyon, which may be one of the most botanically diverse and interesting environments in the entire Santa Catalina range. Cold air flows ("drains") southwest, down from Green Mountain (upper right, elev. 2408 m) and Bear Saddle (elev. 2034 m) into the canyon (elev. 1600-1800 m) essentially along the road over a distance of about 3 km. Water is also concentrated in the canyon and occasional floods have deposited alluvium in the comparatively flat areas near Bear Creek. Soils here can therefore be relatively deep. The creek supports typical riparian species like Arizona sycamore and Arizona (canyon) grape near its banks.

Heading northeast on the Catalina Highway in Bear Canyon. Oak woodland on the left south-facing slope (August). Photo: Steve Smith.

Because of the cooler conditions here, ponderosa and southwestern white pines also occur here at elevations much lower than anywhere else in the Catalinas. Species typically found at lower elevations such as alligator juniper and Mexican blue oak also occur here. 

Bear Creek with very little water present (August). The presence of logs and woody debris suggests that significant flooding has occurred here. Photo: Steve Smith.

Can you visualize the landscape from this topographic map?

Another view of aspect

Here we can see the role of aspect in relation to species composition. The north-facing (northern aspect) slope of Bear Canyon is best categorized as a pine oak forest while the south-facing slope is oak woodland or other more drought-adapted community types.

Pine oak forests in this environment represent a very good example of the fluid nature of plant community boundaries. Oaks are generally adapted to drier and warmer sites while pines are found on those that are somewhat wetter and cooler. The transition areas are known as pine oak forests and there are many patches of this community type in the Catalinas.

Ponderosa pine

Photo: Max Licher.

Another sight identification example: We observe a tree in Bear Canyon with seed-producing cones > 8 cm long with sharp-tipped scales, needles 10-20 cm long, mostly 2-4 per fascicle, and bark yellow to red brown that is deeply furrowed and forms scaly plates.

There are four pines found in the Santa Catalinas. Knowing their appearances, preferred habitats, and elevation ranges would lead us to conclude that this is ponderosa pine. This is the most common pine species in Arizona and is found on open, generally drier slopes between 1500 and 2700 m. Many animals depend on ponderosa pine seeds including squirrels and chipmunks. Because it is so common and mature individuals are tolerant of low-intensity fire, this species is a critical component of natural resource management in many areas of Arizona.

Alligator juniper

In Bear Canyon we also observe a tree that can be up to 10-15 m tall and that based on it's leaves and seed cones is a member of the Cupressaceae. On larger individuals (> 5-8 m tall), the bark is dark gray-brown and cracked into small plates, appearing somewhat like the skin of a reptile. Leaves are scale-like and about 2.5 mm long. Seed cones (lower right) are berry like and are orange brown with a waxy bloom when mature.

Knowing their appearances, preferred habitats, and elevation ranges would lead us to conclude that this is alligator juniper. This species occurs on dry sites at elevations from 750-2700 m in the southwestern United States and northern Mexico.

Piñyon-juniper

A dry woodland higher up

Moving up from Bear Canyon the highway climbs fairly steeply on a south-facing slope up toward a great view of Tucson off to the Southwest at Windy Point.

Then, after turning to the northeast, slopes are now mostly west- and east-facing. Even though we've gone up more than 200 m from Bear Canyon (at Windy Point) these slopes and aspects mean that the plants we see are adapted to more drought-prone environments than we just encountered in Bear Canyon. Here we’ll see mostly plants of the piñyon-juniper community. The two main species here being Mexican piñyon pine and alligator juniper. Given the generally lower density of trees, this is really more of a woodland than a forest.

Piñyon-juniper woodlands are rarely dominated by just these two woody species. In the Catalinas, oaks are very common, such as Emory oak, as are other pines, typically observed at higher elevations.

Evidence of the Bighorn Fire is visible in the upper left of this image taken in August 2020. Here on the edge of the burned area it's possible to see how patchy fire effects can be on large woody species. It's also possible to see dead but still-standing trees (called “snags”) in this image. These were probably killed by the Aspen Fire in 2003, or by drought or bark beetles.

Chaparral

Mexican piñyon pine and alligator juniper are found in relatively dry sites compared to all other conifers in the Santa Catalinas. We will also see point-leaf manzanita here, especially on the steepest south-facing slopes. This is a very striking red-barked shrub that is often a component of a shrub-dominated plant community called chaparral. Chaparral is interspersed with piñyon-juniper and oak woodland communities in the Catalinas. Manzanitas are quite fire-adapted and will often regrow from basal buds that survive fire.

Pointleaf manzanita in the chaparral community in the Santa Catalinas (August). Red bark is characteristic of the species. Photo: Steve Smith.

Pointleaf manzanita flowers (June). Photo: Steve Smith.

Pointleaf manzanita regrowth after a fire from basal buds (October). Photo: Steve Smith.

Pine forest

The first extensive forests we see

The piñyon-juniper and chaparral communities transition quite quickly around Willow Canyon at an elevation of about 2100 m. Here again we enter forest, this time dominated almost entirely by ponderosa pine.

Ponderosa pine. Photo: Steve Smith.

In areas where low-intensity fires have occurred at natural frequencies (about every 10-30 years), ponderosa pine-dominated forests have relatively few, widely spaced, but large, mature trees. In areas where these fires have not occurred regularly, such forests can become much denser and contain many snags. Wildfires and controlled ("prescribed") burns have occurred in many of the coniferous forest communities in the Santa Catalinas. The Bighorn Fire affected this area in 2020.

Careful observation of the forest community can permit description of its recent fire history. To do this, one could look at such things as tree density, range in size of living and dead trees, fire scarring on bark, and presence of burned logs, snags, and species that commonly invade following fires.

A fire-scared living ponderosa pine east of Willow Canyon in the Santa Catalinas (2010). Burned logs and snags are visible in the background. Photo: Steve Smith.

Bear Wallow

Conifers and many broad-leafed species

Bear Wallow is a gently sloping drainage with an ephemeral (flowing for a short time) creek fed by Bear Wallow Spring (elev. 2511 m) and the slopes above it. The creek flows northwest running underneath the highway and then toward Soldier Camp. There it heads west and into Sabino Creek. Given the available moisture and elevation of Bear Wallow (elev. 2400 m), vegetation here is dense with an overstory of coniferous trees and many broad-leafed species such as bigtooth maple, and shrubs like redosier dogwood. Because of the presence of both conifers and broad-leafed species, we would consider this a mixed conifer forest community.

Sufficient moisture and deep soils mean that some very large coniferous trees occur in Bear Wallow. Photo: Kandres Halbrook.

The understory is also diverse, especially in the riparian zone. Golden columbine is one of the most attractive species here. This area was subjected to a variety of water and soil conservation activities during the 20th Century. This involved introducing plants—mostly grasses—not native to the Santa Catalinas and many remain common near the creek. 

Douglas fir

Bear Wallow has numerous large trees, which could be defined as those with stem diameters at breast height (DBH) > 70 cm. Among these large trees, many have thick and corky bark that is deeply fissured and scaly with ridges. The needles, which are not in fascicles, are about 3 cm long. Seed-producing cones have three-pointed bracts between the scales. Given these observations and knowledge of preferred habitats and elevation ranges we would conclude that this is Douglas fir.

Seed cones of Douglas fir are very distinctive. The bracts under the cone's scales are often called “mouse tails” since they resemble the hind legs, tail, and back of a mouse. Bark on younger trees is much smoother and may be reddish brown. The species occurs at elevations between 2000 and 3200 m, but may be lower in some canyons. The tallest trees in Arizona forests are Douglas firs.

Douglas fir with DBH of 60 cm at Bear Wallow. Photo: Steve Smith.

Quaking aspen

Another often tall (> 10 m) tree in Bear Wallow and higher in the Catalinas has smooth whitish/gray bark and nearly round leaves with a sharp point and finely toothed margins. The leaves are shiny green above and dull green beneath with the petiole longer than the blade and flattened.

These observations combined with knowledge of preferred habitats and elevation ranges would lead us to conclude that this is quaking aspen. This species occurs on wet or intermittently wet sites at elevations from 1900 to 3100 m. It is deciduous, often producing vibrant yellow fall color as leaves senesce. Quaking aspen may spread laterally by producing shoots from near-surface roots. Clonal groves develop via this spread after fire or other disturbances. Quaking aspen is the most widely distributed tree in North America, occurring in areas with cold and cool temperate climates from north of the Arctic Circle (near sea level) to near Mexico City (at high elevations) and from coast to coast.

White to gray bark with shallow, horizontal furrows are distinctive of trees in the genus Populus that contains quaking aspen. Photo: Sue Carnahan.

The image at the right was taken in August 2020 after the Bighorn Fire, which burned vegetation on the steep slope in the background. The brown leaves on smaller trees under the aspens in the foreground were desiccated by flame heat, but did not burn.

More evidence of fire

As we proceed up the highway from Bear Wallow toward the village of Summerhaven, evidence of fire is obvious in the vegetation near the road. Broad-leafed multi-stemmed trees, such as Gambel oak and New Mexico locust occur in this image (June 2020) on what were mixed conifer forests with stands dominated by ponderosa pine and Douglas fir before the Aspen Fire in 2003.

Some overstory pines remain, but their density is much lower than before the fire. Snags are still present, seventeen years after the fire.

To the summit of Mt. Lemmon

A much wetter and cooler environment

At the north edge of Summerhaven (elev. 2350 m) we leave the Catalina Highway and head west up East Ski Run Road toward the summit of Mt. Lemmon. This road follows the drainage of Sabino Creek, which is fed by at least four springs in the area. Heading past the Mt. Lemmon Ski Valley, the (now unpaved) road goes up a steep slope to the summit (elev. 2792 m) where the University of Arizona’s Steward Observatory manages a group of telescopes. North of the observatories there are north-facing slopes that have vegetation typical of mixed conifer and quaking aspen forests at much lower elevations (<1000 m) in northern Alberta, Canada, more than 3500 km north of this location. Here we will see many of the plants present at Bear Wallow. However, we may also commonly see corkbark fir, white fir, mountain snowberry, and Rocky Mountain maple, which has more coarsely toothed leaf margins than bigtooth maple.

White fir. Photo: Max Licher

Mt. Lemmon is named for a botanist!

Mt. Lemmon is named for the California botanist and artist  Sara Plummer Lemmon  who explored the Santa Catalina Mountains on her honeymoon in 1881 with her husband John Gill Lemmon and rancher Emerson Stratton. They climbed the north side of the range to the summit approaching from the town of Oracle.

They explored the Southwest in an era when the going was rough. A very interesting  article  based on the Lemmons' letters to friends and family gives us a glimpse of what southern Arizona was like in the 1880s. Many plant species have been named in their  honor .

John and Sara Lemmon. Photo of original by Wynne Brown. Original: Jepson Herbarium, University of California, Berkeley.

Heading home

As we head back, we hope you have a chance to review what we've learned and think about how you might apply this when you are next in the field. For many of you, this next field experience might actually be into the Santa Catalina Mountains. Regardless, our hope is that you may apply this knowledge anywhere.

A quick review

We can be now more aware of the roles that elevation, precipitation, topography, slope, aspect, and fire play in determining the types and abundance of plant species seen.

For example, we know that as elevation increases, precipitation increases and temperature decreases. Cool, moist environments provide suitable habitat for a variety of tree species as we saw at Bear Wallow. In the Santa Catalina Mountains, at a latitude of approximately 32˚N, we have to climb to 2400 m to see this species mix, but they can be found at only 350 m elevation in Portland, Oregon (latitude 45˚N). Thus, in southern Arizona, climbing 2000 m in elevation is equivalent to driving northwest 2000 km.

Here we see Douglas fir, Arizona alder, and white spruce at Bear Wallow. Photo: Steve Smith.

If we overlay our plant communities here, we see that they generally coincide with the elevation, temperature, and precipitation gradients.

However, transitions between plant communities are rarely abrupt. Instead they are influenced by local characteristics of the topography.

For example, in the Catalinas, piñyon-juniper or chaparral communities are found at elevations where one might expect to see pine oak forests. Instead on the southern-western aspects and steep slopes (= less water retention) at these elevations only more drought-tolerant species of the piñyon-juniper or chaparral communities survive.

Similarly, canyons that have a slope with a northern or eastern aspect provide cooler, shady conditions that allow plants typically seen at higher elevations to thrive. We saw this at Bear Canyon where ponderosa pines and other species adapted to higher elevations (cooler, moister environments) were found.

This image was taken on the southeast aspect of Bear Canyon and is looking toward the north-facing slope. The difference between the vegetation on the two slopes (northern vs. southern) is striking and is an example of how multiple factors play a role in determining which species one may find at a particular location.

The photo, at left, was taken at the location shown by the camera icon (August).

Can you visualize the ridges and drainages in this photo? Could you apply this to a contour map?

We learned the importance of maps and orienting oneself to the landscape.

The topographic contour map in the lower half of the image has been rotated to approximate the view shown in the photograph.

We worked through the process of how field botanists sight identify individual species. We saw that this is a integrative process that may use geographical, ecological, and finally botanical information to identify a plant. As you gain experience, more emphasis is placed on characteristics of the plants within any community type. Here is an image of a Sonoran Desert scrub community in Sabino Canyon taken in March. How many apparently different species can you see here?

One final view of the communities we've seen.

Remember to complete the two activities described during the field trip (defining terms in bold text, and describing and photographing an example of an aspect effect on plant growth). Include your responses in your Field Notebook.

Credits

All images not otherwise attributed are the work of Kandres Halbrook or Steven Smith. Other images were graciously donated for educational use from contributors to the Southwestern Environmental Information Network ( SEINet)  consortium of herbaria collection databases.

Vegetation map layers were derived from a database managed by Landscape Fire and Resource Management Planning Tools ( LANDFIRE ), a shared program between the wildland fire management programs of the U.S. Department of Agriculture Forest Service and U.S. Department of the Interior.

Species collection data came from SEINet.

Global 1-arc-second digital elevation data came from the U.S. Geological Survey's Shuttle Radar Topography Mission ( SRTM) . Temperature and precipitation data were obtained from  WorldClim. 

Maps were created using ArcGIS® software by Esri. ArcGIS® and StoryMap™ are the intellectual property of Esri and are used herein under license. Copyright © Esri. All rights reserved. For more information about Esri® software, please visit www.esri.com.

Lower elevation environments (left) are often dominated by cacti such as saguaro and drought-deciduous shrubs such as ocotillo, which is flowering in this image that was taken in March. Less than 45 minutes away by car is Bear Wallow (right), a moist temperate forest whose vegetation (all plants collectively) is more typical of the northern Rocky Mountains than of southern Arizona.

Predicted mean annual precipitation of the Santa Catalina Mountains and the surrounding area based on WorldClim data.

Predicted mean annual temperature of Santa Catalina Mountains and the surrounding area based on WorldClim data.

Image: Google Earth Pro 7.3.3.7786 (August 18, 2018), 3D Scene: ArcGIS® software by Esri

Aspect raster created from a 1-arcsecond digital elevation model

John and Sara Lemmon. Photo of original by Wynne Brown. Original: Jepson Herbarium, University of California, Berkeley.

Game camera image of a desert bighorn sheep on a steep slope in the Santa Catalinas. Photo: Brett Blum.

A paved road runs up Sabino Canyon from a Visitor's Center at the base of the canyon. Trams carry visitors up the canyon stopping at various locations. Lots of folks also walk or ride bikes (at certain times) up and down the canyon. Many trails in the canyon are very popular, especially in cooler periods of the year.

Water flows in Sabino Creek are fed by springs and precipitation over a large watershed on the south side of the Santa Catalinas. Plants found near the creek, in what is called a riparian zone (along the banks of a watercourse), are often very different from those away from this zone.

The creek flows under and sometimes over the road. Flooding and the physical disturbances it causes are an important factor affecting riparian vegetation and the animals associated with it. Photos: Kandres Halbrook.

Typical Sonoran Desert scrub plant community near the base of the Santa Catalinas. The darker green shrubby trees here are velvet mesquite. Photo: Steve Smith.

Two large clumps of fountaingrass, one actively growing (left) and the other appearing to be dormant (August). This image was taken on the downhill (south) side of the Catalina Highway. The rocks occur here because of highway renovation that began in the 1980s. Many weedy species are well adapted to this sort of soil disturbance. Photos: Steve Smith.

The light brown portions of the hillside--elevation 1350 m--are dormant grasses (August). Mature saguaros, although still present, are much less common at this elevation. Photo: Steve Smith.

The Aspen Fire burned for about a month in 2003 over an area of about 34,000 ha and destroyed 340 structures in Summerhaven near the summit of Mt. Lemmon.

October 2016, 13 years after the Aspen fire. Photo: Google.

Blue grama. Photo: Max Licher.

Cheatgrass growing in rocks along roadside drainage of the Catalina Highway (April). Photo: Kandres Halbrook

Cheatgrass along the Catalina Highway near Bear Canyon (April). Photo: Kandres Halbrook.

Heading northeast on the Catalina Highway in Bear Canyon. Oak woodland on the left south-facing slope (August). Photo: Steve Smith.

Bear Creek with very little water present (August). The presence of logs and woody debris suggests that significant flooding has occurred here. Photo: Steve Smith.

Photo: Max Licher.

Pointleaf manzanita in the chaparral community in the Santa Catalinas (August). Red bark is characteristic of the species. Photo: Steve Smith.

Pointleaf manzanita flowers (June). Photo: Steve Smith.

Pointleaf manzanita regrowth after a fire from basal buds (October). Photo: Steve Smith.

Ponderosa pine. Photo: Steve Smith.

A fire-scared living ponderosa pine east of Willow Canyon in the Santa Catalinas (2010). Burned logs and snags are visible in the background. Photo: Steve Smith.

Sufficient moisture and deep soils mean that some very large coniferous trees occur in Bear Wallow. Photo: Kandres Halbrook.

Douglas fir with DBH of 60 cm at Bear Wallow. Photo: Steve Smith.

White to gray bark with shallow, horizontal furrows are distinctive of trees in the genus Populus that contains quaking aspen. Photo: Sue Carnahan.

White fir. Photo: Max Licher

Here we see Douglas fir, Arizona alder, and white spruce at Bear Wallow. Photo: Steve Smith.

The photo, at left, was taken at the location shown by the camera icon (August).